LEICESTER, England (AP) - He was king of England, but for centuries he lay without shroud or coffin in an unknown grave, and his name became a byword for villainy.
On Monday, scientists announced they had rescued the remains of Richard III from anonymity - and the monarch's fans hope a revival of his reputation will soon follow.
In a dramatically orchestrated press conference, a team of archaeologists, geneticists, genealogists and other scientists from the University of Leicester announced that tests had proven what they scarcely dared to hope - a scarred and broken skeleton unearthed under a drab municipal parking lot was that of the 15th-century king, the last English monarch to die in battle.
Jo Appleby, who led the exhumation of the remains found during a dig at a Leicester parking lot, speaks Monday about the conclusion that the remains are those of England’s King Richard III, missing for 500 years.
Lead archaeologist Richard Butler said that a battery of tests proved "beyond reasonable doubt" that the remains were the king's.
Lin Foxhall, head of the university's school of archaeology, said the discovery "could end up rewriting a little bit of history in a big way."
Few monarchs have seen their reputations decline as much after death as Richard III. He ruled England between 1483 and 1485, during the decades-long battle over the throne known as the Wars of the Roses, which pitted two wings of the ruling Plantagenet dynasty - York and Lancaster - against one another.
His brief reign saw liberal reforms, including the introduction of the right to bail and the lifting of restrictions on books and printing presses.
But his rule was challenged, and he was defeated and killed by the army of Henry Tudor, who took the throne as King Henry VII and ended the Plantagenet line. Britain's current monarch, Queen Elizabeth II, is distantly related to Richard, but is not a descendant.
After his death, historians writing under the victorious Tudors comprehensively trashed Richard's reputation, accusing him of myriad crimes - most famously, the murder of his two nephews, the "Princes in the Tower."
William Shakespeare indelibly depicted Richard as a hunchbacked usurper who left a trail of bodies on his way to the throne before dying in battle, shouting "My kingdom for a horse."
The location of Richard's body was unknown for centuries. He died in August 1485 at the Battle of Bosworth Field in the English Midlands, and records say he was buried by the Franciscan monks of Grey Friars at their church in Leicester, 100 miles north of London.
The church was closed and dismantled after King Henry VIII dissolved the monasteries in 1538, and its location eventually was forgotten by most local residents.
There were tales that the king's bones had been dug up and thrown in a nearby river in the 16th century.
Then last year a team led by University of Leicester archaeologist Richard Buckley identified a possible location of the grave through map regression analysis, starting with a current map of the general area of the former church and analyzing earlier maps to discover what had changed and not changed. Ground-penetrating radar was used to find the best places to start digging.
The team began excavating in a parking lot last August. Within a week they had located thick walls and the remains of tiled floors. Soon after, they found human remains - the skeleton of an adult male who appeared to have died in battle.
He had been buried unceremoniously, with no coffin or shroud - plausible for a despised and defeated enemy.
Archaeological bone specialist Jo Appleby, a lecturer in human bioarchaeology at Leicester, said study of the bones provided "a highly convincing case for identification of Richard III."
Appleby said the 10 injuries to the body were inflicted by weapons such as swords, daggers and halberds and were consistent with accounts of Richard being struck down in battle - his helmet knocked from his head - before his body was stripped naked and flung over the back of a horse in disgrace.
The remains also displayed signs of scoliosis, a form of spinal curvature, consistent with contemporary accounts of Richard's appearance.